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Ataenius Harold 1867
Ataenius Harold 1867: 100.

 
Ataenius chilensis (Solier)
Photo by Paul Skelley.
Classification
Superfamily.....
Family...........
Subfamily......
Tribe.............
Scarabaeoidea
Scarabaeidae
Aphodiinae
Eupariini
 

 

Aphodiinae Overview

Aphodiinae Key

Eupariini Key
 
 
 
 
 
 
 
 
 
 
 
 
 

Type species. Ataenius scutellaris Harold 1867, by indirect original designation (monotypy).

 
 

Diagnosis. Body length 1.5-7.0 mm, elongate, parallel-sided or not, usually reddish-brown to black, few with color patterns, some dorsally setose, others distinctly agrillaceous. Head usually convex, surface smooth, granulate, wrinkled, or rugose. Clypeus with distinct median emargination, toothed or not on either side, margin usually thickened at middle. Pronotum variable shape, usually not explanate; often with setal fringe. Mesosternum carinate between coxae. Mesotarsi usually not longer than tibia. Abdominal fluting distinct. External sexual dimorphisms subtle, if present.

 
 
 
 
 
 
 
 
 

Distribution. Worldwide.

 

Composition. Worldwide, Ataenius contains over 300 species, 189 of which occur in the New World (Stebnicka 2007c).
[MAYBE PUT THIS LIST OF SPECIES ON A SEPARATE WEB PAGE, so genus account would be small. Number of species in New World actually 189, but some species below have yet to be officially moved to other genera. UPDATE ranges based on Stebnicka 2007c).]
Ataenius abancay Stebnicka 2005: 106-107, Peru.
Ataenius abdominalis Petrovitz 1973: 154, Brazil.
Ataenius aciculus Hinton 1937: 187, Haiti, Dom. Rep.
                  = Ataenius apicecoloratus Balthasar 1941: 168.
Ataenius aequalis Harold 1880: 40, N-Argentina to S- US, and West Indies.
                  = Ataenius insulicola Chapin 1940: 28, St. Vincent.
                  = Ataenius titschacki Balthasar 1941: 167, Argentina.
Ataenius aequatorialis Petrovitz 1961: 355, Galapagos.
                  = Ataenius aequinoctialis Petrovitz 1961: 355 (lapsus calami).
Ataenius alternatus (Melsheimer 1844: 137) [Oxyomus], SE-US, Mexico.
Ataenius apicalis Hinton 1937: 195, Honduras, Mexico, E-USA.
Ataenius arenosus Harold 1868: 86, Brazil.
Ataenius arrowi Hinton 1936: 415, Galapagos.
                  = Ataenius leleupi Petrovitz 1976: 286.
Ataenius atramentarius (Erichson 1847: 110) [Euparia], Galapagos (introduced), Peru, Ecuador.
                  = Ataenius dampieri Petrovitz 1961: 356.Ataenius attenuator Harold 1874: 22,                   Colombia, Venezuela W.Ind. (St.Lucia, St Vincent, Trinidad)
                  = Ataenius abditoides Chapin 1940: 18.
                  = Ataenius denticulatus Petrovitz 1965: 176.
Ataenius abditus (Haldeman 1848: 106) [Aphodius (Oxyomus)] , widespread New World.
Ataenius balthasari Petrovitz 1973: 163-165, Brazil.
Ataenius barberi Cartwright 1974: 51, TX,AZ-USA.
Ataenius beattyi Chapin 1940: 17, W.Indies. (Guadeloupe, Virgin Islands, Jamaica, DR, PR, Barbados).
                  = Ataenius cameneni Chalumeau and Gruner 1974: 801.
Ataenius bicolor Petrovitz 1963: 642, Jamaica, Cuba.
Ataenius bispinulosus Schmidt 1911: 52, Argentina, Paraguay.
Ataenius blapoides Balthasar 1947: 51, Brazil, Paraguay.
Ataenius bolivarensis Stebnicka 2007a: 73, Venezuela, French Guiana.
Ataenius bolivari Stebnicka 2001: 269, Venezuela.
Ataenius borgmeieri Hinton 1936: 472, Mexico.
Ataenius borjae Stebnicka 2005: 107-108, Bolivia.
Ataenius brevicollis (Wollaston 1854: 229) [Oxyomus], SE-USA, West Indies (Bahamas, Cuba, DR, Guadeloupe), Madeira Is., Canary Is., Portugal.
                  = Pleurophorus chopardi Paulian 1937: 75.
                  = Ataenius frankenbergeri Balthasar 1938: 56.
                  = Ataenius lherminiari Paulian 1947: 43.
                  = Auperia sulcatula Chevrolat 1864: 413.
Ataenius brevis Fall 1930: 98, E-USA.
Ataenius brunneus Schmidt 1922: 102. Argentina.
Ataenius buenavistae Stebnicka 2001: 282, Bolivia, Peru.
Ataenius caicarae Stebnicka 2005: 108-109, Venezuela.
Ataenius californicus Horn 1887: 84, SW-USA.
Ataenius canoasus Stebnicka 2007a: 76, Brazil, Argentina.
Ataenius carinatipennis Petrovitz 1973: 151-152, Brazil.
                  = Ataenius granulipennis Petrovitz 1973: 152-154.
Ataenius carinator Harold 1874: 20, Mexico to S. America, FL-USA (introduced), West Indies.
                  = Ataenius vincentiae Arrow 1903: 513.
Ataenius cartago Stebnicka 2007a: 55, Costa Rica, Mexico, Guatemala.
Ataenius cartwrighti Chalumeau and Gruner 1974: 813, Guadeloupe.
Ataenius castaniellus Bates 1887: 95 (see Galante et al. 2003) Mexico, Guatemala, Honduras, El Salvador.
                  = Ataenius liogaster var. castaniellus Bates 1887: 95.
Ataenius catarinaensis Stebnicka 2007a: 76, Brazil, Paraguay.
Ataenius catenulatus (Erichson 1847: 110) [Euparia], Peru.
Ataenius cavagnaroi Cartwright 1970: 55, Galapagos.
Ataenius chapini Hinton 1937: 3, Mexico to Panama.
                  = Ataenius frankorum Deloya 2000: 55.
Ataenius chinacotae Stebnicka 2007a: 64, Colombia, Bolivia, Costa Rica, Ecuador, Panama.
Ataenius chilensis (Solier 1851: 72) [Aphodius], Chile.
                  = Ataenius modestus Schmidt 1912: 199.
Ataenius chocabambae Stebnicka 2005: 112-113, Bolivia.
Ataenius clavatus Schmidt 1916: 103, Brazil, Argentina.
Ataenius clitellarius Petrovitz 1973: 165, Brazil.
Ataenius cognatus (LeConte 1858: 65) [Euparia], SW-USA, Mexico.
Ataenius columbicus Harold 1880: 39, Argentina, Brazil, Colombia, Paraguay, Venezuela.
                  = Ataenius fastus Petrovitz 1970: 236.
                  = Ataenius laterigranosus Balthasar 1947: 61.
                  = Ataenius tesari Balthasar 1947: 50.
Ataenius communis Hinton 1936: 421, Mexico to S.America.
Ataenius complicatus Harold 1869: 102, Mexico to Argentina.
                  = Ataenius loretti Martinez 1952: 104.
Ataenius confertus Fall 1909: 162, SW-USA, Baja California.
                  = Ataenius cribratus VanDyke 1928: 156.
Ataenius convexus Robinson 1940: 149, TX-USA.
Ataenius corrosus Chapin 1940: 19, Cuba.
Ataenius costulifer Balthasar 1951: 354[340], Peru.
Ataenius crenator Harold 1876: 97, Brazil.
                  = Ataenius nigritus Petrovitz 1973: 171.
Ataenius crenatostriatus (Blanchard 1843: 184) [Oxyomus], Bolivia, Argentina.
Ataenius crenaticollis Petrovitz 1973: 158, Brazil, Peru.
Ataenius crenulatus Schmidt 1910: 359, Mexico to Argentina, West Indies.
                  = Ataenius rickardsi Hinton 1938: 123.
                  = Ataenius rickardasi [sic] in Deloya 1994: 52.
                  = Ataenius ricardi [sic] in Galante, Stebnicka and Verdú 2003:                                     291.                                   
Ataenius cribrithorax Bates 1887: 95, Mexico to Panama, W. Indies.
Ataenius cristobalensis Cook and Peck 2000: 290, Galapagos.
Ataenius cucutae Stebnicka 2007a: 75, Colombia, Ecuador.
Ataenius cylindrus Horn 1871: 289, SE-USA.
                  = Ataenius cylindricus Horn in Schmidt 1922: 454 (misspelling).
                  = Ataenius horni Harold 1874: 19.
Ataenius depilis Petrovitz 1976: 284, Ecuador, Venezuela, Trinidad.
Ataenius desertus Horn 1871: 289, SW-USA, Mexico.
Ataenius ecruensis Stenicka and Lago 2005: 65-66, E-USA.
Ataenius elegans Harold 1868: 85, Brazil, Venezuela.
Ataenius elisaensis Stebnicka 2002: 270.
Ataenius elongatus (Palisot de Beauvois 1811: 104) [Scarabaeus], Hispaniola.
Ataenius erratus Fall 1930: 96, SE-USA.
Ataenius exiguus Brown 1932: 10, SE-USA.
Ataenius fattigi Cartwright 1948: 151-153, E-USA.
Ataenius floreanae Cook and Peck 2000: 292, Galapagos.
Ataenius forsteri Balthasar 1960: 4, S. America.
Ataenius glabriventris Schmidt 1911: 52, Mexico, Central America to Venezuela.
Ataenius glaseri Cartwright 1974: 57, E-USA.
Ataenius gracilis (Melsheimer 1844: 137) [Oxyomus (Pleurophorus)], US to South America, West Indies, Micronesia, Pacific Islands.
                  = Saprosites nocturnus Nomura 1943: 77.
Ataenius griffini Cartwright 1974: 77, TX-USA.
Ataenius gruneri Chalumeau 1979: 229, Cuba.
Ataenius guanacastae Stebnicka 2005: 115-116, Costa Rica, Panama.
Ataenius guayasi Stebnicka 2001: 270, Ecuador.
Ataeniius guriensis Stebnicka 2005: 117, Venezuela..
Ataenius heinekeni (Wollaston 1854: 228) [Oxyomus], Cuba, SE-US to Central Amer.
                  = Auperia rhyticephalus Chevrolat 1864: 413.
                  = Ataenius floridanus Brown 1930: 3.                 
                  = Ataenius solitarius Blatchley 1928: 69.
                  = Ataenius vexator Harold 1869: 103.
Ataenius hesperius Cartwright 1974: 61, W-USA.
Ataenius hirsutus Horn 1871: 228, SW-USA, Mexico.
Ataenius hispaniolae Chalumeau 1982: 325, Dominican Republic.
Ataenius hispidus Harold 1867: 83, Venezuela, Suriname.
Ataenius holopubescens Hinton 1938: 124-126, Mexico, Guatemala.
Ataenius howdeni Chalumeau 1978: 51, Antigua.
Ataenius huanus Stebnicka 2007a: 68, Peru, Ecuador.
Ataenius hygrophilus Paulian 1947: 39, Guadeloupe.
Ataenius imbricatoides Schmidt 1909: 36, Argentina, Brazil.
Ataenius imbricatus (Melsheimer 1844: 136) [Aphodius], Central and E-USA to Argentina, W.Indies.
                  = Ataenius sordidus Harold 1869: 103.
Ataenius impiger Schmidt 1916: 104, southern S.America.
                  = Ataenius latrigranulatus Balthasar 1941: 166-167.
                  = Ataenius perpunctatus Balthasar 1961: 124.
Ataenius impressus (Petrovitz 1963: 643) [Dialytes], Brazil, Bolivia.
Ataenius incanus Balthasar 1951: 340, Peru.
Ataenius inquisitus Horn 1887: 81, SW-USA.
Ataenius insculptus Horn 1887: 70, SE-USA.
                  = Ataenius sculptilis LeConte 1878: 402 [not Ataenius sculptilis Harold 1868:                   86].
Ataenius insulae Chalumeau and Gruner 1974: 805, Guadeloupe.
Ataenius iquitosae Stebnicka 2007a: 52, Peru.
Ataenius isabelae Franz 1985: 97, Galapagos.
Ataenius jamaicensis Chapin 1940: 33, Jamaica.
Ataenius jardinensis Stebnicka 2002: 278, Cuba.
Ataenius jelineki Chalumeau 1982: 323, Haiti.
Ataenius klapperichi Howden 1978: 378, Dominican Republic.
Ataenius kochi Balthasar 1941: 167, Brazil.
Ataenius koelleri Balthasar 1963: 288, Paraguay to Panama.
Ataenius lamarensis Stebnicka 2007a: 70, Peru, Bolivia, Argentina.
Ataenius lanei Petrovitz 1973: 157, Brazil.
Ataenius languidus Schmidt 1910: 31, S- USA to Venezuela, West Indies.
                  = Ataenius linelli Cartwright 1944: 28.
Ataenius latus Petrovitz 1963: 316, Venezuela.
Ataenius lenkoi Petrovitz 1973: 177, Brazil.
Ataenius liogaster Bates 1887: 94, Mexico to N. South Amer., West Indies, Galapagos, Australia, Micronesia.
                  = Ataenius edwardsi Chapin 1940: 26-27, W.Indies (not Cuba).
                  = Ataenius hoguei Cartwright and Spangler 1981: 785-789, Mexico.
                  = Ataenius kelatianus Balthasar 1965: 316.
                  = Ataenius nitidulus Nomura 1943: 78.
                  = Ataenius orbicularis Schmidt 1914: 697.
Ataenius lobatus Horn 1871: 287, SW-USA, Baja California-Mexico.
                  = Ataenius laeviventris Horn 1887: 74.
Ataenius londrinae Stebnicka 2007a: 74, Brazil, Bolivia, Argentina, Paraguay, Ecuador.
Ataenius longiclavus Petrovitz 1970: 233, Brazil.
Ataenius lucanus Horn 1871: 288, Baja California.
Ataenius luctuosus (Burmeister 1877: 411) [Euparia], Argentina, Paraguay.
                  = Ataenius tectus Petrovitz 1973: 175-176.
Ataenius luteomargo Chapin 1940: 36, W.Ind. (Lesser Antilles, PR), Panama, Venezuela, Suriname.
                  = Ataenius terminalis Arrow 1903: 512, [not Ataenius terminalis Chevrolat                   1864: 414].
                  = Ataenius versicolor Hinton 1937: 183, [not Ataenius versicolor Schmidt                   1916: 105].
Ataenius martinezi Petrovitz 1973: 162, Argentina.
Ataenius miamii Cartwright 1934: 200, E-USA, Bahamas.
Ataenius michelii Chalumeau 1978: 49, Puerto Rico.
Ataenius montanus Schmidt 1911: 32, Ecuador.
                  = Ataenius grossepunctatus Petrovitz 1976: 288.
Ataenius morator Harold 1869: 103, Brazil, Argentina, to W.Indies.
                  = Ataenius insitivus Balthasar 1961: 125.
                  = Ataenius picipes Fleutiaux and Salle 1890[1889]: 397.
                  = Ataenius tenebrosus Arrow 1903: 512.
Ataenius nanus Degeer 1774: 318, Suriname.
Ataenius napoensis Stebnicka 2007a: 65, Ecuador, Peru, Venezuela.
Ataenius noronhai Stebnicka 2007a: 55, Brazil (Island Fernando de Noronha).
Ataenius noques Stebnicka 2007a: 77, Argentina, Brazil, Paraguay.
Ataenius nugator Harold 1880: 41, Central and S. America.
                  = Ataenius bolivianus Hinton 1937: 189.
                  = Ataenius duplex Hinton 1936: 418.
Ataenius oaxacaensis Stenicka and Lago 2005: 78-79, Mexico.
Ataenius ocumarensis Stebnicka 2001: 279, Venezuela.
Ataenius oklahomensis Brown 1930: 4, S.Central USA (AK, LA, OK, TX).
Ataenius onkonensis Stebnicka 2005: 124, Ecuador.
Ataenius opacipennis Schmidt 1910: 360, Brazil, Suriname, Paraguay, Venezuela.
                  = Ataenius drifti Endrodi 1963: 50.
Ataenius opatrinus Harold 1867: 82, Central and S.America [FL-USA ?].

                  = Ataenius narialis Petrovitz 1973: 173.
                  = Ataenius paraperforatus Deloya and Ibanez-Bernal 2000: 318.
                  = Ataenius woodruffi Cartwright 1974: 72.
Ataenius opatroides (Balthasar 1847: 185) [Oxyomus], Uruguay, Argentina, Chile.
Ataenius ovatulus Horn 1871: 286, SE-USA (MD-IL and south).
                  = Ataenius lecontei Harold 1874: 20.
Ataenius pacificus (Sharp 1879: 90) [Aphodius], Hawaii, Tahiti.
Ataenius palmaritoensis Stebnicka 2007a: 72, Venezuela, Colombia, Peru, Ecuador.
Ataenius panamensis Hinton 1936: 425, Panama.
Ataenius parallelipennis Petrovitz 1973: 173, Guatemala.
Ataenius pearlensis Stebnicka 2006: 97-98, Panama.
Ataenius peregrinator Harold 1877: 96, Indonesia, Philippines, Micronesia, Vietnam, USA (FL, AZ) to Colombia and Ecuador.
                  = Ataenius jalapensis Bates 1887: 99-100 (See Galante et al. 2003).
                  = Ataenius polyglyptus var. jalapensis Bates 1887: 100.
                  = Ataenius polyglyptus var. hieronymi Bates 1887: 100.
                  = Ataenius nunenmacheri Cartwright 1974: 53.
                  = Ataenius stroheckeri Cartwright 1974: 54.                 
                  = Ataenius sumatrensis Balthasar 1941: 184.
Ataenius pereirai Petrovitz 1970: 237, Brazil.
Ataenius perforatus Harold 1867: 98, Mexico to S. America.
Ataenius pertuga Balthasar 1961: 125, Brazil.
Ataenius petrovitzi Balthasar 1960: 6, Ecuador to Argentina to Brazil.
                  = Ataenius auropunctatus Petrovitz 1973: 166.
Ataenius picinus Harold 1867: 281, nearly cosmopolitan (Americas,Australia N.Zeal.,N.Caled., Fiji, Vanuatu).
                  = Ataenius alegrus Balthasar 1947: 50.
                  = Ataenius boucomontii Paulian 1937: 41.
                  = Ataenius darlingtoni Hinton 1937: 179.
                  = Ataenius duplopunctatus Lea 1923: 6.
                  = Ataenius paracognatus Balthasar 1961: 123.
                  = Ataenius queirosii Paulian 1934: 219.
                  = Saprosites rugosus Richards 1959: 41.
                  = Ataenius salutator Fall 1930: 99.
Ataenius platensis (Blanchard 1846: 185) [Oxyomus], Argentina to S-USA, W. Indies.
                  = Ataenius anticus Fall 1930: 105.
                  = Ataenius degallieri Chalumeau 1990: 340.
                  = Ataenius granchacoensis Balthasar 1939: 28.
                  = Ataenius heyrovskyi Balthasar 1960: 5.
                  = Ataenius histrionicus Balthasar 1947: 49.
                  = Ataenius integer Harold 1868: 86, S-USA.
Ataenius plaumanni Petrovitz 1973: 160, Brazil, Argentina.
Ataenius polyglyptus Bates 1887: 99, West Indies, Panama, Guatemala.
                  = Ataenius polyglyptus var. intermedius Bates 1887: 100.
                  = Ataenius ecuadoriensis Petorvitz 1970: 235.
Ataenius pseudocarinator Balthasar 1947: 54, French Guiana, USA.
                  = Ataenius vandykei Cartwright 1974: 39.
Ataenius pseudoclavatus Stebnicka 2005: 129, Argentina.
Ataenius pseudocommunis Stebnicka 2001: 267, Venezuela.
Ataenius pseudostercorator Stebnicka 2003: 236-237, Bolivia.
Ataenius pseudousingeri Galante, Stebnicka and Verdú 2003: 295, Mexico, Guatemala..
Ataenius punctatohirsutus Schmidt 1909: 36, Argentina.
Ataenius punctipennis Harold 1868: 86, S-America, Trinidad, Grenada, Carriacou,
                  = Ataenius subopacus Chapin 1940: 24..
Ataenius purator Harold 1868: 85, widespread South America.
                  = Ataenius gothi Balthasar 1933: 9.
                  = Ataenius gagates Petrovitz 1963: 317.
                  = Ataenius splendens Endrodi 1963: 52-54.
Ataenius quintanaroo Stebnicka 2006: 102, Mexico.
Ataenius raccurti Chalumeau 1978: 50, Dominican Rep.
Ataenius robustus Horn 1871: 285, Central-USA.
Ataenius rubrotessellatus (Blanchard 1843: 185) [Oxyomus], Bolivia.
Ataenius sabinoi Cartwright 1974: 38, AZ-USA, Baja California-Mexico.
Ataenius saltae Stebnicka 2007a: 71, Argentina, Bolivia.
Ataenius santarosae Stebnicka 2007a: 78, Peru, Argentina, Bolivia.
Ataenius saramari Cartwright 1939: 360, FL-USA.
Ataenius saulensis Stebnicka 2006: 109-111, French Guiana.
Ataenius scabrelloides Petrovitz 1962: 131, Costa Rica to Bolivia to Brazil, USA (FL).
                  = Ataenius superficialis Cartwright 1974: 30-31.
Ataenius scabrellus Schmidt 1909: 118, Venezuela, West Indies to south Florida-USA.
                  = Ataenius havanensis Balthasar 1938: 56.
                  = Ataenius fleutiauxi Paulian 1947: 42.
Ataenius scalptifrons Bates 1887: 100, Mexico and Central America.
Ataenius schmidti Stebnicka 2003: 240, Argentina, Paraguay, Brazil.
                  = Ataenius agrillaceous Schmidt 1916: 103, not A. agrillaceous Chevrolat                   1869.
Ataenius sculptilis Harold 1868: 86, S. America, West Indies.
Ataenius sculptor Harold 1868: 85, SW-USA to Colombia.
                  = Ataenius oblongus Horn 1871: 286.
Ataenius scutellaris Harold 1867: 82, Mexico to Brazil, W.Ind, Bahamas, Madagascar, Vanuatu, Seychelles, Mascarenes, Malaya.
                  = Ataenius auberti Paulian 1937: 42.
                  = Ataenius frater Arrow 1903: 512.
Ataenius semipilosus VanDyke 1928: 158, SW-USA.
Ataenius setiger Bates 1887: 98, Mexico, SW-USA.
                  = Ataenius pseudohirsutus Cartwright 1974: 37, SW-USA.
Ataenius siminasus Petrovitz 1973: 155-156, Brazil, Peru.
Ataenius skelleyi Stebnicka 2007a: 54, Brazil, Venezuela.
Ataenius spretulus (Haldeman 1848: 106) [Aphodius], SE-Canada, USA.
                  = Ataenius consors Fall 1930: 104 [not Ataenius consors Blackburn 1904:
                  161].
                  = Ataenius falli Hinton 1934: 119.
Ataenius steinheili Harold 1874: 18, Mexico, Colombia, Grenada, Trinidad, Venezuela.
                  = Ataenius grenadensis Chalumeau 1982: 327, Grenada.
Ataenius stephani Cartwright 1974: 81, AZ-USA, Mexico.
Ataenius stercorator (Fabricius 1775: 20) [Scarabaeus], S.America (S.of equator).
                  = Ataenius opacus Harold 1867: 100.
Ataenius strigatus (Say 1823: 212) [Aphodius], SE-Canada, USA, Mexico.
Ataenius strigicauda Bates 1887: 96, Mexico to Argentina, W.Indies, Madeira.
                  = Ataenius aspericollis Petrovitz 1973: 178.

Ataenius talpoides Stebnicka 2001: 279, Argentina.
                  = Ataenius talpa Balthasar 1947: 52.
Ataenius tambopatae Stebnicka 2001: 280, Peru.
Ataenius tarumensis Stebnicka 2007a: 63, Brazil.
Ataenius temperei Chalumeau and Gruner 1974: 799, W.Ind. (Dominica, Guadeloupe).
Ataenius terminalis (Chevrolat 1864: 414), Cuba, Jamaica.
                  = Auperia ciliata Chevrolat 1864: 414.
Ataenius texanus Harold 1874: 23, Central USA, Mexico, Central America and West Indies.
                  = Ataenius punctifrons Cartwright 1974: 59.
Ataenius tovarensis Stebnicka 2001: 268, Venezuela.
Ataenius tuberculatus Schmidt 1911: 15, Colombia to Brazil and Bolivia.
Ataenius usingeri Hinton 1937: 191, Mexico to Panama..
Ataenius utahensis Cartwright 1974: 61, UT-USA.
Ataenius variopunctatus Schmidt 1922: 102, Argentina.
Ataenius versicolor Schmidt 1916: 105, Hispaniola.
                  = Ataenius haitianus Hinton 1937: 184.
Ataenius vinacoensis Stebnicka 2006: 105, Argentina.
Ataenius waltherhorni Balthasar 1938: 55, Cuba, E-USA.
Ataenius wenzeli Horn 1887: 77, SE-USA.
                  = Ataenius ludovicianus Fall 1930: 100.
                  = Ataenius rudellus Fall 1930: 103, FL-USA.
Ataenius yungasus Stebnicka 2001: 275, Bolivia.

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
Life History. The majority of Ataenius appear to be detritivores in various niches; organic accumulations along margins of open sands, leaf litter, ant or packrat nests, old mammal dung, etc. One, A. holopubescens, is known to occur in detritus piles of leaf-cutter ants, Atta spp. (Stebnicka 2007b).  Most specimens in collections came to lights, others were sifted out of various substrates. Details on the biology of most remain unknown, those with some studies include: Cruz and Martinez (2002, A. apicalis and A. sculptor), Martinez and Cruz (2002, A. apicalis and A. sculptor), Wegner and Niemcyzk (1981, A. spretulus). Ataenius spretulus is also known as the “black turfgrass ataenius” and is reported as a turf pest in many publications. A few species have been transported nearly worldwide with human activity, including A. brevicollis, A. liogaster, A. picinus, etc.
 
 
 
 
 
 
 
 
 
 
 
 
 
Larvae. The larvae of A. brevis, A. erratus, A. imbricatus, A. opatrinus, A. ovatulus, A. picinus, A. platensis, and A. strigicauda have been described or diagnosed in Jerath (1960) and Verdú and Galante (1999).
 
 
 

References:
Cartwright, O. L. 1974. Ataenius, Aphotaenius, and Pseudataenius of the United States and Canada. Smithsonian Contributions to Zoology 154: 1-106.
Cruz‑R., M. and I. Martinez‑M. 2002. Data on nesting and preimaginal development in two Mexican species of Ataenius Harold, 1867 (Coleoptera, Scarabaeoidea, Aphodiidae; Eupariinae). Folia Entomologica Mexicana 41(1): 1‑5.
Jerath, M. L. 1960. Notes on larvae of nine genera of Aphodiinae in the United States. Proceedings of the United States National Museum, Washington 111(3425): 43-94.
Martinez-M., I., and R. Cruz-R. 2002. Fenologia y ciclos reproductivos en Ataenius apicalis Hinton y A. sculptor Harold (Coleoptera: Aphodiidae). Bulletin de la Société Entomologique de France 107: 177-186.
Smith, A. B. T., and P. E. Skelley. 2007. A review of the Aphodiinae (Coleoptera: Scarabaeidae) of southern South America. Zootaxa 1458: 1-80. [http: //www.museum.unl.edu/research/entomology/platycoeliams.htm]
Stebnicka, Z. T. 2001. The New World species of Ataenius Harold, 1867. I. Revision of the A. crenator-group, A. nugator-group and A. perforatus-group (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia 44: 253-283.
Stebnicka, Z. T. 2002. The New World species of Ataenius Harold, 1867. II. Revision of the West Indian A. terminalis-group (Coleoptera: Scarabaeidae: Eupariini). Acta Zoologica Cracoviensia 45: 259-281.
Stebnicka, Z. T. 2003. The New World species of Ataenius Harold, 1867. III. Revision of the A. imbricatus-group sensu lato (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia 46(3): 219-249.
Stebnicka, Z. T. 2004. The New World species of Ataenius Harold, 1867. IV. Revision of the A. strigicauda-group (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia, 47(3-4): 211-228.
Stebnicka, Z. T., and P. K. Lago. 2005. The New World species of Ataenius Harold, 1867. V. Revision of the A. strigatus-group (Scarabaeidae: Aphodiinae: Eupariini). Insecta Mundi 19(1-2): 55-83.
Stebnicka, Z. T. 2005. The New World species of Ataenius Harold, 1867. VI. Revision of the A. aequalis-platensis-group (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia, Invertebrata 48B: 99-138.
Stebnicka, Z. T. 2006. The New World species of Ataenius Harold, 1867. VII. Revision of the A. complicatus-group (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia, Invertebrata 49B: 89-114.
Stebnicka, Z. T. 2007a. The New World species of Ataenius Harold, 1867. VIII. Revision of the A. scutellaris-group and the A. texanus-carinator group with descriptions of new species (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia, Invertebrata 50B: 45-81.
Stebnicka, Z. T. 2007b. New species of Iarupea Martínez and morphological specializations among related taxa associated with ants and termites (Coleoptera: Scarabaeidae: Eupariini). Revue Suisse de Zoologie 114(3): 573-590.
Stebnicka, Z. T. 2007c. The genus Ataenius Harold, 1867 (Coleoptera: Scarabaeidae) of the world. Iconography. Institute of Systematics and Evolution of Animals, Polish Academy of Science, Krakow, Poland. 155 pp.  
Verdú, J. R., and E. Galante. 1999. Larvae of Ataenius (Coleoptera: Scarabaeidae: Aphodiinae): generic characteristics and species Descriptions. European Journal of Entomology 96(1): 57‑68.
Wegner, G. S., and H. D. Niemczyk. 1981. Bionomics and phenology of Ataenius spretulus. Annals of the Entomological Society of America 74(4): 374‑384.

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
   
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